“Recruitment in a social carnivore before and after harvest” is the title of a recent paper published on the Journal Animal Conservation of the Zoological Society of London that shows collateral impacts of harvesting wolves:
“We attributed just 18–38% of pup mortality directly to harvest and suggest that there are indirect effects of harvest on recruitment that may be associated with changes in group size and structure. Models that do not include both direct and indirect effects of harvest on recruitment may underestimate the potential impact of harvest on population growth in social species.”
The article does not have open access and it is available HERE but you can read it in this LINK.
Other recommended reads: http://www.klamathconservation.org/science_blog/conservation/?p=192